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Bible Encyclopedias
Ear
1911 Encyclopedia Britannica
or (common Teut.; O.E. Bare, Ger. Ohr, Du. oor, akin to Lat. auris, Gr. as), in anatomy, the organ of hearing. The human ear is divided into three parts - external, middle and internal. The external ear consists of the pinna and the external auditory meatus. The pinna is composed of a yellow fibro-cartilaginous framework covered by skin, and has an external and an internal or cranial surface. Round the margin of the external surface in its upper three quarters is a rim called the helix (fig. 1, a), in which is often seen a little prominence FIG. - The Ear as seen in Section.
n, External auditory meatus.
o, Membrana tympani.
p, Tympanum.
points to malleus.
2, to incus.
3, to stapes.
4, to cochlea.
5, 6, 7, the three semicircular canals.
8 and 9, facial and auditory nerves.
known as Darwin's tubercle, representing the folded-over apex of a prick-eared ancestor. Concentric with the helix and nearer the meatus is the antihelix (b), which, above, divides into two limbs to enclose the triangular fossa of the antihelix. Between the helix and the antihelix is the fossa of the helix. In front of the antihelix is the deep fossa known as the concha (fig. 1, d), and from the anterior part of this the meatus passes inward into the skull. Overlapping the meatus from in front is a flap called the tragus, and below and behind this is another smaller flap, the antitragus. The lower part of the pinna is the lobule (e), which contains no cartilage. On the cranial surface of the pinna elevations correspond to the concha and to the fossae a, Helix.
b, Antitragus.
c, Antihelix.
d, Concha.
e, Lobule.
f, Mastoid process.
g, Portio dura.
h, Styloid process.
k, Internal carotid artery. 1, Eustachian tube.
m, Tip of petrous process.
of the helix and antihelix. The pinna can be slightly moved by the anterior, superior and posterior auricular muscles, and in addition to these there are four small intrinsic muscles on the external surface, known as the helicis major and minor, the tragicus and the antitragicus, and two on the internal surface called the obliquus and transversus. The external auditory meatus (fig. r, n) is a tube running at first forward and upward, then a little backward and then forward and slightly downward; of course all the time it is also running inward until the tympanic membrane is reached. The tube is about an inch long, its outer third being cartilaginous and its inner two-thirds bony. It is lined by skin in its whole length, the sweat glands of which are modified to secrete the wax or cerumen.
The middle ear or tympanum (fig. i, p) is a small cavity in the temporal bone, the shape of which may perhaps be realized by imagining a hock bottle subjected to lateral pressure in such a way that its circular section becomes triangular, the base of the triangle being above. The neck of the bottle, also laterally compressed, will represent the Eustachian tube (fig. i, 1), which runs forward, inward and downward, to open into the nasopharynx, and so admits air into the tympanum. The bottom of the bottle will represent the posterior wall of the tympanum, from the upper part of which an opening leads backward into the mastoid antrum and so into the air-cells of the mastoid process. Lower down is a little pyramid which transmits the stapedius muscle, and at the base of this is a small opening known as the iter chordae posterius, for the chorda tympani to come through from the facial nerve. The roof is formed by a very thin plate of bone, called the tegmen tympani, which separates the cavity from the middle fossa of the skull. Below the roof the upper part of the tympanum is somewhat constricted off from the rest, and to this part the term "attic" is often applied. The floor is a mere groove formed by the meeting of the external and internal walls. The outer wall is largely occupied by the tympanic membrane (fig. i, o), which entirely separates the middle ear from the external auditory meatus; it is circular, and so placed that it slopes from above, downward and inward, and from behind, forward and inward. Externally it is lined by skin, internally by mucous membrane, while between the two is a firm fibrous membrane, convex inward about its centre to form the umbo. Just in front of the membrane on the outer wall is the Glaserian fissure leading to the glenoid cavity, and close to this is the canal of Huguier for the chorda tympani nerve. The inner wall shows a promontory caused by the cochlea and grooved by thetympanic plexus of nerves; above and behind it is the fenestra ovalis, while below and behind the fenestra rotunda is seen, closed by a membrane. Curving round, above and behind the promontory and fenestrae, is a ridge caused by the aqueductus Fallopii or canal for the facial nerve. The whole tympanum is about half an inch from before backward, and half an inch high, and is spanned from side to side by three small bones, of which the malleus (fig. i, 1) is the most external. This is attached by its handle to the umbo of the tympanic membrane, while its head lies in the attic and articulates posteriorly with the upper part of the next bone or incus (fig. i, 2). The long process of the incus runs downward and ends in a little knob called the os orbiculare, which is jointed on to the stapes or stirrup bone (fig. i, The two branches of the stapes are anterior and posterior, while the footplate fits into the fenestra ovalis and is bound to it by a membrane. It will thus be seen that the stapes lies nearly at right angles to the long process of the incus. From the front of the malleus a slender process projects forward into the Glaserian fissure, while from the back of the incus the posterior process is directed backward and is attached to the posterior wall of the tympanum. These two processes form a fulcrum by which the lever action of the malleus and incus is brought about, so that when the handle of the malleus is pushed in by the membrane the head moves out; the top of the incus, attached to it, also moves out, and the os orbiculare moves in, and so the stapes is pressed into the fenestra ovalis. The stapedius and tensor tympanic muscles, the latter of which enters the tympanum in a canal just above the Eustachian tube to be attached to the malleus, modify the movements of the ossicles.
The mucous membrane lining the tympanum is continuous through the Eustachian tube with that of the naso-pharynx, and is reflected on to the ossicles, muscles and chorda tympani nerve. It is ciliated except where it covers the membrana. tympani, ossicles and promontory; here it is stratified.
The internal ear or labyrinth consists of a bony and a membranous part, the latter of which is contained in the former. The bony labyrinth is composed of the vestibule, the semicircular canals and the cochlea. The vestibule lies just internal to the posterior part of the tympanum, and there would be a communication between the two, through the fenestra ovalis, were it not that the foot plate of the stapes blocks the Dc way. The inner wall of the vestibule is separated from the bottom of the internal auditory meatus by a plate of bone pierced by many foramina for branches of the auditory nerve (fig. i, 9), while at the lower part is the opening of the aqueductus vestibuli, by means of which a communication is established with the posterior cranial fossa. Posteriorly the three semicircular canals open into the vestibule; of these the external (fig. i, 7) has two independent openings, but the superior and posterior (fig. i, 5 and 6) join together at one end and so have a common opening, while at their other ends they open separately. The three canals have therefore five openings into the vestibule instead of six. One end of each canal is dilated to form its ampulla. The superior semicircular canal is vertical, and the two pillars of its arch are nearly external and internal; the external canal is horizontal, its two pillars being anterior and posterior, while the convexity of the arch of the posterior canal is backward and its two pillars are superior and inferior. Anteriorly the vestibule leads into the cochlea (fig. z, 4), which is twisted two and a half times round a central pillar called the modiolus, the whole cochlea forming a rounded cone something like the shell of a snail though it is only about 5 mm. from base to apex. Projecting from the modiolus is a horizontal plate which runs round it from base to apex like a spiral staircase; this is known as the lamina spiralis, and it stretches nearly half-way across the canal of the cochlea. At the summit it ends in a little hook named the hamulus. The modiolus is pierced by canals which transmit branches of the auditory nerve to the lamina spiralis.
The membranous labyrinth lies in the bony labyrinth, but does not fill it; between the two is the fluid called perilymph, while inside the membranous labyrinth is the endolymph. In the bony vestibule lie two membranous bags, the saccule (fig. 2, S) in front, and the utric]e (fig. 2, U) behind; each of these has a special patch or macula to which twigs of the auditory nerve are supplied, and in the mucous membrane of which specialized hair cells are found (fig. 3, p). Attached to the maculae are crystals of carbonate of lime called otoconia. The membranous semicircular canals are very much smaller in section than the bony; in the ampulla of each is a ridge, the crista acustica, which is covered by a mucous FIG. 2. - Diagram of the Membranous Labyrinth.
DC, Ductus cochlearis.
dr, Ductus reuniens.
S, Sacculus.
U, Utriculus.
dv, Ductus endolymphaticus. SC, Semicircular canals.
(After Waldeyer.) FIG. 3. - cl, Columnar cells covering the crista acustica; p, peripheral, and c, central processes of auditory cells; n, nerve fibres. (After Riidinger.) membrane containing sensory hair cells like those in the maculae. All the canals open into the utricle. From the lower part of the saccule a small canal called the ductus endolymphaticus (fig. 2, dv) runs into the aqueductus vestibuli; it is soon joined by a small duct from the utricle, and ends, close to the dura mater of the posterior fossa of the cranium, as the saccus endolymphaticus, which may have minute perforations through which the endolymph can pass. Anteriorly the saccule communicates with the membranous cochlea or scala media by a short ductus reuniens (fig. 2, dr). A section through each turn of the cochlea shows the bony lamina spiralis, already noticed, which is continued right across the canal by the basilar membrane (fig. 4, bm), thus cutting the canal into an upper and lower half and connected with the outer wall by the strong spiral ligament (fig. 4, sl). Near the free end of the lamina spiralis another membrane called the membrane of Reissner (fig. 4, mR) is attached, and runs outward and upward to the outer wall, taking a triangular slice out of the upper half of the section. There are now three canals seen in section, the upper of which is the scala vestibuli (fig. 4, SV), the middle and outer the scala media, ductus cochlearis or true membranous cochlea (fig. 4, DC), while the lower is the scala tympani (fig. 4, ST). The scala vestibuli and scala tympani communicate at the apex of the cochlea by an opening known as the helicotrema, so that the perilymph can here pass from one canal to the other. At the base of the cochlea the FIG. 4. - Transverse Section through the Tube of the Cochlea. Modiolus.
0, Outer wall of cochlea.
SV, Scala vestibuli. ST, Scala tympani.
DC, Ductus cochlearis. mR, Membrane of Reissner.
perilymph in the scala vestibuli is continuous with that in the vestibule, but that in the scala tympani bathes the inner surface of the membrane stretched across the fenestra rotunda, and also communicates with the subarachnoid space through the aqueductus cochleae, which opens into the posterior cranial fossa. The scala media containing endolymph communicates, as has been shown, with the saccule through the canalis reuniens, while, at the apex of the cochlea, it ends in a blind extremity of consider able morphological interest called the lagena.
The scala media contains the essential organ of hearing or organ of Corti (fig. 4, oc), which lies upon the inner part of the basilar membrane; it consists of a tunnel bounded on each side of the inner and outer rods of Corti; on each side of these are the inner and outer hair cells, between the latter of which are found the supporting cells of Deiters. Most externally are the large cells of Hensen. A delicate membrane called the lamina reticularis covers the top of all these, and is pierced by the hairs of the hair cells, while above this is the loose membrana tectoria attached to the periosteum of the lamina spiralis, near its tip, internally, and possibly to some of Deiter's cells externally. The cochlear branch of the auditory nerve enters the lamina spiralis, where a spiral ganglion (fig. 4, sg) is developed on it; after this it is distributed to the inner and outer hair cells.
For further details see Text-Book of Anatomy, edited by D. J. Cunningham (Edinburgh, 1906); Quain's Elements of Anatomy (London, 1893); Gray's Anatomy (London, 1905); A Treatise on Anatomy, edited by H. Morris (London, 1902); A Text-Book of Human Anatomy, by A. Macalister (London, 1889).
